Debunking Tradition:
Charters Rewrite Hiltoun Pedigree.

With the launch of new ‘Research & Resource’ pages for Guthrie of Eassie and Guthrie of Hiltoun, it’s time to revisit one of the most persistent ancestral lines in Angus genealogy—and why primary charters now tell a different story.

Genealogy enthusiasts know the frustration all too well: we inherit beautifully compiled family trees that have been copied, reprinted, and trusted for generations—only to discover they rest on shaky foundations. Traditional pedigrees, even those published in the 19th century, were often built from secondary summaries rather than original charters, retours, or sasines. The people who compiled them faced the same obstacles we do today: conflated names, missing dates, and the ever-present temptation to connect dots that don’t quite align. Sometimes all it takes is one overlooked document to turn “settled fact” into a fascinating question mark.

Bottom line: The traditional pedigree makes John Guthrie, 1st of Hiltoun, a younger son of Sir Alexander Guthrie of Guthrie and Kincaldrum. The surviving primary records now point elsewhere. By 1544, John Guthrie was already recognized as heir of David Guthrie of Eassie, and the 1545/1546 Hiltoun records appear to represent an additional grant to an established Eassie landholder rather than an inheritance from the senior Guthrie line.

The Traditional Pedigree – A 150-Year Assumption

For nearly 150 years, since Andrew Jervise’s ‘Epitaphs & Inscriptions (1879) and Alexander J Warden‘s first volume of Angus or Forfarshire (1880), John Guthrie, 1st of Hiltoun (also styled of Hilton or Hilltoun), has been placed as a younger son of Sir Alexander Guthrie, 2nd Baron of Guthrie and 3rd of Kincaldrum (d. 1513 at Flodden) and his wife Hon. Margaret Lyon, a daughter of Lord Glamis.

Several children are credited to the couple:

• David Guthrie, the eldest son who d. 1513 at Flodden by whom the Guthrie of that Ilk line continued via his son, Andrew Guthrie, 3rd of Guthrie.
• Alexander Guthrie, who inherited the title and lands of Kincaldrum, splitting the line into two branches.
• John Guthrie, a younger son who inherited Hiltoun
• Modern pedigrees also list George Guthrie of Kincreich, and a daughter, Elizabeth Guthrie.

This made Hiltoun a direct late cadet of the senior Guthrie line — a virtually uncontested story that has been repeated time and again in Burke’s Landed Gentry, Stirnet, and countless family trees.

If you descend from the Guthrie of Hiltoun and Guthrie of Craigie lineages, these are the things you need to know about the origins of your ancestors.

The 1495 Bombshell and the Compressed Timeline

Sometimes, all it takes is one new fact to open your eyes to other connections. Before we get to John Guthrie, 1st of Hiltoun, we need to focus our attention on Margaret Lyon of Glamis whose marriage contract creates a serious chronological problem for the traditional pedigree.

No doubt you have read that the wife of Sir Alexander Guthrie, 2nd of Guthrie, 3rd of Kincaldrum, was Margaret Lyon, the daughter of John Lyon, 3rd Lord Glamis, and Elizabeth Scrymgeour. She was one of seven daughters (Violetta, Janet, Christian, Agnes, Margaret, Mariota, and Elizabeth), all of whom have spouses identified in The Scots Peerage.

Except–there is no mention of a Guthrie marriage in the records of Glamis Castle.

Even Jervise and Warden could not name her.

“Sir Alexander [Guthrie] married a daughter of Lord Glamis.” — Alexander J. Warden, Angus or Forfarshire, Vol. III, p. 389

The specific identification of Alexander’s wife as Margaret Lyon appears consistently only in later 19th- and 20th-century secondary sources (Burke’s Landed Gentry, Stirnet, etc.). The Lyon of Glamis connection has remained especially appealing to Guthrie descendants because any daughter in that line serves as a gateway ancestor to royal Stewart descent.

Daughters of John Lyon, 3rd Lord Glamis and Elizabeth Scrymgeour:

No other Margaret Lyon from any branch of the Lyon family (Glamis main line or the Cossins/Wester Ogil cadets) appears in contemporary or near-contemporary records at the right age to have been Alexander Guthrie’s wife.

There are two vital “anchor dates” to consider in this Guthrie family: 1495 and 1513.

• On 10 June 1495, Margaret Lyon entered a formal marriage contract with James Rynd, younger of Broxmouth (tocher (dowry): 400 merks). The original document still rests at Glamis Castle and is explicitly cited in The Scots Peerage. [Transcript of Original Scan]

• On 9 September 1513, Sir Alexander Guthrie and his son, David Guthrie, were both killed at the Battle of Flodden.

No primary marriage contract or charter has ever been found naming her in connection with Alexander Guthrie. If he married a Lyon daughter (Full Table in Reference Section), the marriage contract has not been indexed.

A marriage to Margaret Lyon is still definitely possible, but it would have to be after 1495 when she was contractually available for marriage. No primary documents have been found to indicate that James Rynd inherited Broxmouth, or that he had children by Margaret Lyon. If James Rynd died soon after 1495, or their contract of marriage was never solemnized, a Guthrie-Lyon marriage could have taken place between 1495-1513.

Any children for the couple would have to be born between 1495 and 1513–roughly a window of 18 years. This still seems plausible until those birth year estimates and current pedigrees show their children born between the 1460s – 1490s.

By September 1513 the following had supposedly occurred:

• Eldest son David Guthrie was old enough to fight and die at Flodden and already had a son, Andrew Guthrie, who was old enough to succeed to the main Guthrie estates immediately after the battle.
• Both Alexander Guthrie and his son David fell together at the Battle of Flodden on 9 September 1513. There is no single official list of Flodden casualties, but contemporary legal records confirm their deaths. Warden notes that Alexander “surrounded by friends and vassals, attended his sovereign to the fatal field of Flodden, and there fell, with his eldest, David, his three brothers-in-law, David, William, and George Lyon, and his nephew, Sir Thomas Maule.”
• Just a few months later, on 15 May 1514, an Instrument of Sasine was granted in favour of Andrew Guthre as (grand)son & heir of the deceased Alexander Guthre of Guthre, kt., of the lands of Guthre. (In 16th-century Scottish feudal practice a minor heir’s sasine normally named a tutor or curator; none is mentioned here.)

If Margaret Lyon was Alexander Guthrie’s only wife and the mother of any children, this timeline is highly improbable. Even if their eldest son, David Guthrie, was old enough to fight at the Battle of Flodden, his son Andrew would likely only be an infant rather than being considered old enough to receive title over the lands and barony of Guthrie the following year. Room is also required for the younger Alexander Guthrie to inherit the title and lands of Kincaldrum. John Guthrie is in place as a younger son who was said to inherit Hiltoun either before or shortly after his father’s death in 1513.

Our search into the Traditional Pedigree of John Guthrie 1st of Hiltoun has highlighted a problem with the identification of his mother. Yet there has been nothing shown to shake up his listed father, Sir Alexander Guthrie, 2nd of Guthrie, 3rd of Kincaldrum.

Before dismissing the traditional pedigree outright, we have to ask whether Hiltoun could have been granted to a younger son during Sir Alexander’s lifetime. Such grants were possible—but the surviving record does not show such a grant to John.

The lands of Guthrie and Kincaldrum were distributed before Sir Alexander Guthrie’s 1513 death. He had already granted the Kincaldrum lands to his younger son Alexander by charter dated 23 August 1508 (RMS ii, 3259), a common practice for younger sons during the father’s lifetime.

Hiltoun seems to have been a division of the Guthrie lands, but there was no division or grant to any John Guthrie specifically documented as a son of Sir Alexander Guthrie of Guthrie & Kincaldrum either before or in the thirty year period after his death.

• On 15 Oct 1545, a Precept of Clare Constat by Alexander Guthrie, 4th of Guthrie, to John Guthrie of Hilton of Guthrie. (NLS, Saltoun Papers, Ch.14167)

• On 10 May 1546, A Charter and Instrument of Sasine by Alexander Guthrie (4th) of Guthrie, son of Andrew Guthrie (3rd) of Guthrie, to John Guthrie of Esse of parts of Hilton and Easttown of Guthrie. (NLS, Saltoun Papers, Ch.14168–14169)

Importantly, the documents reveal that John Guthrie was styled “of Esse” prior to his acquisition of Hiltoun. There is one other key record:

• Eleven months earlier, on 3 November 1544, a Precept of Clare Constat by James, Lord Ogilvy had already recognized John Guthrie “as heir of deceased David Guthrie of Esse” in the lands of Kirktoun of Esse (NRS GF16/3/42). This confirms that John was already an established landholder “of Esse” before receiving the Hiltoun grant from Alexander Guthrie, 4th of Guthrie.

This is not evidence that John was simply a “son or relative” being handed land as a new vassal. It shows he was already a recognized, landed heir in the Eassie line (under Ogilvy superiority) by late 1544. The 1545/46 Hiltoun grant was therefore an additional infeftment of an established local laird — not the first time John was being “set up” with property.

The Primary Evidence Continues

There are only a few surviving documents mentioning the Guthries of Eassie, either with or without a connection to Hiltoun. Based on the date of the first record in 1454, we can surmise that the John Guthery of Esse was not the same man as John Guthrie of Eassie & Hiltoun from the 1540s, since he would already have been of legal age in 1454. He could be the same John Guthrie from the 1490 or 1493 documents. We also have two sets of father-son combinations: John and son Alexander (1493) and David and son John (1544). These are a great place to speculate on the formation of John Guthrie of Eassie & Hiltoun’s ancestral pedigree.

Pedigree Possibilities for John Guthrie of Eassie/Hiltoun

That 1544 document solidifies John Guthrie of Eassie & Hiltoun’s parentage. He was the son of David Guthrie of Eassie, not the son of Sir Alexander Guthrie and Margaret Lyon.

These documents were not the first time “Guthrie of Eassie” has come up. The Guthrie1 Pedigree on Stirnet assigns “William Guthrie of Eassie” as a son of Alexander Guthrie, 1st of Kincaldrum, and his wife Marjory Guthrie. No primary records have been found so far identifying him in connection to that family, but it does create speculation as to whether Eassie was a cadet branch of the Kincaldrum line.

Kincaldrum, Eassie, and Hiltoun occupy the same compact corner of central Angus. Ordnance Survey / Google Maps data show:

• Kincaldrum → Eassie: ≈ 5 miles (8 km) (Kincaldrum House ruins lie just northwest of Gateside on the A90; Eassie village/church is west of Glamis on the A94.)
• Kincaldrum → Hiltoun (Hilton of Guthrie): ≈ 9–10 miles (14–16 km) (Hiltoun lies north of Guthrie Castle, roughly 7 miles east of Forfar.)
• Eassie → Hiltoun: ≈ 8 miles (13 km)

Proximity invites the likelihood of relatedness. If Alexander Guthrie, 2nd of Guthrie, 3rd of Kincaldrum, is not John Guthrie of Eassie & Hiltoun’s father, perhaps the Eassie line is tied to the family in another way. If William Guthrie of Eassie turns out to be a legitimate option, the Eassie line would be a Cadet Branch of Kincaldrum.

The Guthrie DNA Project has revealed the presence of multiple unique genetic lines in Angus, which may require us to consider the possibility that the Guthries of Eassie, Hiltoun, and by extension the Guthries of Craigie originate from their own individual family group. If this is the case, the earliest documented Guthrie of Eassie is the John Guthrie from the 1454 inquest.

• Traditional Kincaldrum to Guthrie to Hiltoun line has been ruled out for John Guthrie.
  • John Guthrie of Hiltoun is not presently proven as a son of Sir Alexander Guthrie of Guthrie/Kincaldrum.
  • The 1544 Eassie record identifies him instead as heir of David Guthrie of Eassie.
  • Any Kincaldrum connection must now be sought through the earlier Eassie line, not through the traditional Sir Alexander → John of Hiltoun descent.
  • If Alexander Guthrie, 2nd of Guthrie, 3rd of Kincaldrum, did have a son named John Guthrie, he does not appear to be connected to the lands of Eassie & Hiltoun as the 1544 document specifically named “John Guthrie of Esse ” as their son & heir of “David Guthrie of Esse.”
• Eassie Cadet via Kincaldrum: In this scenario, John Guthrie descends from Alexander Guthrie, 1st of Kincaldrum, through the speculative son William of Eassie. The first verifiable John of Eassie appears too early to be the same man who acquired Hiltoun in 1545/46. Outcome: Connected to the Kincaldrum branch, but bypasses the main Guthrie of that Ilk line that begins with David 1st of Guthrie / 2nd of Kincaldrum. So, John would descend from Kincaldrum & Eassie, but not Guthrie of that Ilk.

• Independent Eassie Branch: The Eassie line has no proven link to William or the Kincaldrum Guthries as there are really no primary documents to support it. The earliest documented John of Eassie could represent a completely separate Guthrie family that later acquired Hiltoun. People tend to forget that there were other living Guthries prior to Alexander Guthrie, 1st of Kincaldrum. Any connection to the broader Kincaldrum / Guthrie of that Ilk tree is now uncertain.

Why This Matters – Methodology and Ripple Effects

This correction does not diminish the Guthrie of Hiltoun or Craigie lines. It strengthens them by placing them where the surviving records actually lead. The traditional pedigree made John Guthrie of Hiltoun a younger son of the senior Guthrie/Kincaldrum line, but the primary evidence identifies him first as John Guthrie of Eassie, heir of David Guthrie of Eassie. Whether that Eassie family ultimately proves to be a cadet branch of Kincaldrum or a separate Angus Guthrie family remains an open research question. What can no longer be treated as settled fact is the direct descent from Sir Alexander Guthrie and Margaret Lyon.

Continued Research Needed

Further Clarity May be Found in:

• Full transcriptions of the Glamis Castle contract and any later Lyon-Guthrie instruments.
• Closer examination of the Eassie cadet line (possible William of Eassie link).
• Full transcriptions of the primary Eassie / Hiltoun documents.
• Clarification of George of Kincreich’s exact parentage.

We welcome your input! If you hold transcriptions from the Register of the Great Seal, Arbroath Abbey records, or unpublished Glamis material, please share in the comments or email the blog. The Eassie and Hiltoun pages will be updated as new evidence surfaces.

Read the full research:
→ Guthrie of Eassie – Resources & Research
→ Guthrie of Hiltoun – Resources & Research

Primary / Archival Catalogue Sources

1. DOCUMENT: Records of the Guthrie Family of Guthrie, Angus. National Library of Scotland, Ref: GD188.
2. DOCUMENT: Inquest Member: “John Guthery of Esse” was one of 15 “noble and worthy men” selected to settle property claims in the region. The Carnegie-Kinnaird inquisition of 1454 occurred because the Carnegie family charters for Kinnaird castle had been destroyed by fire after the Battle of Brechin in 1452. The inquest established Carnegie territorial rights in the region. Fraser, William. History of the Carnegies, Earls of Southesk, and of their Kindred, Vol. 1, p.18; Contemporary transcript of Brief and Retour at Kinnaird. Digital Repository: Internet Archive.
3. National Records of Scotland, Ref: GD16/3/144. Charter by William Fentoune of Cukiscardane to Sir James Evilly of Irony, kt. Witnesses include: John Guthrie of Essy. 31 Aug 1481.
4. National Records of Scotland, Ref: GD121/3/20. Papers in action over lands of Balmier, 1485-1511, 1573. Letters of execution by Alexander Lyndesay of Ochtirmunsy, sheriff depute of Forfar, at command of royal letters upon possession of certain annuals in town of Kyremur pertaining to Archibald, earl of Anguss, for sum of eight score merks (4 merks 8 d. less) obtained upon said earl by Thomas Fothringam of Powry, narrating that said Thomas caused summon David Ogilvy of that ilk, Alexander Lovell of Ballumby, John Dempstar of Caraldstone, William Maxwell of Tellin, Wat Ramsay of Dunnone, Alexander Fentoun of Ogyll, William Fentoun of Kirkistoun, Hew Watterstoune of that ilk, John Guthre of Esse, John ?Seres of Cragnathro, Alexander Strathachin of Brigtoun, David Narne of the Baky, and Andrew Neve of Mathy, who priced superiority and annuals of tenement and 3 acres of land of John Crab’s, extending to 30s yearly, and superiority with annual of tenement and 2 acres of land of John Fettas, extending yearly to 20s, and superiority with annual of tenement and 3 acres of land of David Alane extending yearly to 15s annual, together with superiority with annual of tenement and other half acre of land of sir John Skynner’s extending yearly to 15s., together with superiority and annual of an acre of land and tenement of Robert Butyr’s extending yearly to 10s., lying in town of Kyremuir, 20 June 1486. Traces of 6 applied seals in dorso.
5. National Records of Scotland, Ref: GD/121/3/27, p.3. Writs of Lands of Brighty. A special retour before Andrew, lord Grey, sheriff of Forfar, in favour of Nicholas Fothryngham as heir of deceased Thomas Fotheryngham, his father, in lands of Westerbrychti and Westerpoury and others, 30 Oct 1490. There were 15 members of the inquest among them John Guthrie of Esse, David Ogilvy of that Ilk and Alexander Guthre of that Ilk. Other key witnesses for this discussion: James Rynde of Broxsmouth, and James Reynde, grandson (nepos) of said James. This is 1493, just 2 years before the marriage contract of James Rynd, younger of Broxmouth, and Margaret Lyon of Glamis.
6. National Records of Scotland, Ref: SP13/45. Warrant for remission under the Great Seal in favour of Robert Maull of Panmure, Thomas Maull, John Guthre of Colytstoun, Thomas Guthre, David Ramsay, David Air, John Guthre of Esse, Hercules Guthre in Lowrane, and Robert Rollok, they having left the party of insurrection. At Edinburgh. Signed by James, Earl of Arran, governor, and John, Abbot of Paisley, Keeper of the Privy Seal, with postscript in holograph of said John that if the above persons joined in an insurrection or refused assistance when required then above warrant was to be of no avail. Jan 1543-1544.
7. National Records of Scotland, Ref: GD16/3/42. Precept of clare constat by James, Lord Ogilvy, to John Guthrie as heir of deceased David Guthrie of Esse, in the lands of Kirktoun of Esse. 3 Nov. 1544
8. National Library of Scotland, Saltoun Papers, Ch.14167 Precept of clare constat by Alexander Guthrie of Guthrie to John Guthrie of Hilton, of Guthrie, 15 October 1545. [Transcript from Digital Scan]
9. National Library of Scotland, Saltoun Papers, Ch.14168, Charter by Alexander Guthrie, son of Andrew Guthrie of Guthrie, to John Guthrie of Esse of parts of Hilton and Easttown of Guthrie., 10 May 1546. [Transcript from Digital Scan]
10. National Library of Scotland, Saltoun Papers, Ch.14169, Instrument of sasine to John Guthrie of Esse of parts of Hilton and Easttown of Guthrie, 10 May 1546. 
11. National Library of Scotland, Saltoun Papers, Ch.14177, Instrument of renunciation by James Betoun of Melgo and Lutres Betoun his wife of an annualrent from Hilton of Guthrie, 11 April 1580. 
12. National Library of Scotland, Saltoun Papers, Ch.14185, Instrument of resignation by Alexander Guthrie of Guthrie in favour of Mr Walter Lindsay of Kemphill, of Hilton of Guthrie and superiority of Langlands, 15 November 1583. 
13. National Library of Scotland, Saltoun Papers, Ch.14194, Charge at instance of Mr Patrick Lindsay… prebendary of Hilton and Langlands, 30 November 1588. 
14. National Library of Scotland, Saltoun Papers, Legal Instruments. Instrument of renunciation by James Wishart… of half of Hilton of Guthrie, 5 May 1589. 
15. National Library of Scotland, Saltoun Papers, Ch.14196, Assignation by Alexander Wishart of Carnbege and James Wishart his son to Gabriel Guthrie… of the wadset of Hilton of Guthrie, 10 March 1589/90. 
16. National Library of Scotland, Saltoun Papers, Ch.14280, Contract between Sir Walter Lindsay of Balgavies and David Lindsay, his son, and Alexander Guthrie of Guthrie and Alexander Guthrie, his son, concerning Hilton and Langlands, 19 July 1603. 
17. National Library of Scotland, Saltoun Papers, Ch.14288, Contract between David Lindsay of Balgays and David Guthrie, son of David Guthrie of Hilton, 8 February 1614. 
18. National Library of Scotland, Saltoun Papers, Ch.14153–14211, collection description noting later documents concerning Guthrie of Guthrie and the lands of Hilton and Langlands, acquired by Lindsay of Balgays in 1603. 
19. Scotlands People. National Records of Scotland, 1779, John Guthrie of Guthrie (Coats of Arms Volume 1 Page No. 519Z, page 1 of 1).

• This is the matriculation of arms of John Guthrie of Guthrie Chief of that ancient surname eldest son an heir of John Guthrie of Guthrie by Joan daughter of the Reverend Mr James Hodge of Bathkennar, Minister of the Gospel.
  • Per the numbering system, this document applies to John Guthrie, 16th of Guthrie, who was the product of multiple Guthrie lines. His 3xGGF was John Guthrie, Bishop of Moray, a direct paternal line descendant of John Guthrie of Eassie & Hiltoun. The bishop purchased the lands and barony of Guthrie making him 11th of Guthrie. His daughter Bathiah Guthrie was married to her cousin Francis Guthrie 3rd of Gagie, whose descent goes through the Kincaldrum to Guthrie to Gagie line.
  • The relevance of this 1779 document is that it lists the genealogy of John Guthrie, 16th of Guthrie, including the Guthrie-Lyon connection. This may be Warden’s source. This document names Margaret Lyon as the wife of Sir Alexander Guthrie, although it mistakenly identified her father as the 5th Lord Glammis instead of the 3rd. Excerpt: “Andrew was son and heir of David Guthrie apparent heir of Guthrie who was killed with his Father and three uncles by his mother together with his cousin Sir Thomas Maule of Panmure at the Battle of Floundoun (Flodden) the ninth of September 1513 and (blank) eldest daughter of Forquhard MacIntosh of MacIntosh by Giles second daughter of High second Lord Lovet which David was son and apparent heir of Sir Alexander Guthrie of Guthrie and Margaret daughter of John fifth Lord Glammis which Sir Alexander was so and heir of Sir David Guthrie of Guthrie Chief of his name by Janet daughter of Sir Archibald Dundas of Dundas Knight…. etc, etc.”

Secondary Genealogical / Compiled Sources

1. John Anderson, ed., Calendar of the Laing Charters, A.D. 854–1837, Belonging to the University of Edinburgh(Edinburgh: James Thin, 1899), 54, no. 214, Internet Archive, accessed May 24, 2026
   
   214. 18th May 1493.] Instrument [much torn] narrating that James [Stewart], Earl of Buchan and lord of Auchirhous, gave sasine to Alexander Guthrie, son and apparent heir of Patrick Guthrie of Halcartoune, of the lands of Hakartoune with pertinents, which were resigned into the hands of the said Eail, hy the said Patrick, who reserved his own liferent and the terce due to Janet Guthrie, his spouse. Done on the lands, 18th May 1493. Witnesses, James Rynde of Broxsmoutht, George Crechtoune, James Rynde, grandson (nepos) of said James, William Petra, John Wynsister, John Guthrie of Essy and Alexander his son, John Kylgour and David Mortimar, chaplains. John of Caigill, presbyter of St. Andrews diocese, by imperial authority notary public. [49, Box 1.
2. Warden, Alexander J. (1882) Angus or Forfarshire: The Land and People, Descriptive and Historical. Vol. III. Dundee: Charles Alexander & Co.
3. Paul, James Balfour, ed. The Scots Peerage. Vol. VIII. Lyon of Glamis family, pp.272-278. Edinburgh: David Douglas, 1911.
4. Burke’s Landed Gentry (majority of editions), placing John Guthrie, 1st of Hiltoun, as a son of Sir Alexander Guthrie and Margaret Lyon.
   • Burke, Bernard (1850) [Burke’s Gentry] Vol. 1: A genealogical and heraldic dictionary of the landed gentry of Great Britain & Ireland, for 1850L comprising particulars of upwards of 100.,000 individuals by John B. Burke. See: Guthrie of Guthrie. Digital Repository: Internet Archive.
   • Burke, Bernard, Sir. (1882). A genealogical and heraldic history of the landed gentry of Great Britain & Ireland. 6th ed. with supplement. London: Harrison. Digital Repository: Internet Archive
   • Burke’s Landed Gentry of Great Britain, 19th ed., Vol. 1, The Kingdom in Scotland, ISBN 0-9711966-0-5. The edition reportedly places John Guthrie of Hiltoun as fourth son of Sir David Guthrie 1st of Guthrie, 2nd of Kincaldrum, a placement that conflicts with other pedigree versions and creates chronological difficulty.
5. Jervise, Andrew (1853) History and Traditions of the Land of the Lindsays in Angus and Mearns. Rewritten and corrected by James Gimmick. Edinburgh: Sutherland and Knox. Digital Repository: Internet Archive.
6. Jervise, Andrew (1879) Epitaphs & Inscriptions from Burial Grounds & Old Buildings in the North East of Scotland with Historical, Biographical, Genealogical, and Antiquarian Notes, also an Appendix of Illustrative Papers, by the late Andrew Jervise, F.S.A. Scot., VOL. II. with a memoir of the author. Edinburgh: David Douglas. Digital Repository: Internet Archive.
   • p.149: Sir David was Justiciary of Scotland in 1473, with a fee of £ 100 Scots (Treasurer’.s Accts., i. 68). He died in 1474, and had two legitimate children, a son and a daughter. The latter became the wife of Sir Thomas Maule of Panmure, and the former, Sir Alexander, who bought the lands of Ballindean, 1468-9, married a daughter of Lord Glamis, by whom he had one daughter and four sons. The youngest son of this marriage was ancestor of John Guthrie of Hilton, from whom the Bishop of Moray was descended.
7. Stirnet Genealogy Database – Guthrie01 Pedigree.
8. WEBSITE: The Peerage – “John Guthrie, 1st of Hilton“John
9. WIKITREE: John Guthrie 1st aka of Hiltoun